Unexpected error when registering using Autofill

A previous event registrant is signing up for a new event using the Autofill link and encounters an Unexpected Error screen.
Please Chat with Support and reference this article. 

In your chat, please include the following:
*url address of the error page
*event URL
*Supporter ID and username of the participant(s) receiving the error message

Environment

 Blackbaud Sphere

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